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ŠUMARSKI LIST 13/2005 str. 154     <-- 154 -->        PDF

S. Kucbel: TI Ili STRUCTURE AND NATURAL REGENERATION OF A SUBALP1NF. SPRUCE FOREST ... Šumarski list- SUPLEMFNT (2005). 144-153
The analyses of natural regeneration was conducted cies and height classes (up to 20 cm, 21-50 cm, 51-80
on the transect. All individuals according to tree spe- cm, 81-130 cm, from 131 cm to dbh 1 cm) were noted.


RESULTS


Stand structure


The values of basic dendrometric traits on particular
research plots are summarized in Table 1. On the
basis of the obtained diameter frequencies and regarding
other stand characteristics it is possible to consider
the stem number of 1,000-1,500 ha"1 as sufficient
(all living individuals with dbh over 1 cm). If only the
large timber is taken into consideration (individuals
with dbh over 7 cm), the stem number should range
from 500 to 1,000 ha"´. The stem numbers with less
than 500 ha"1 can be one of the indicators of an unstable
homogenous structure or of the presence of wider parts
without continuous forest stand.


Diameter distribution curves are one of the basic
characteristics of a stand structure (Fig. 1). On every
observed research plot the distribution curve has more
or less distinctly decreasing shape, which is typical for
a differentiated structure (very close to a selection forest
structure). Some visible differences between particular
research plots follow from the figures. Although
the distribution on the PRP 1 has a decreasing shape,
there is a quite high stem number in higher diameter
classes (trees of the upper layer). The regulation of the
structure should head to the reduction of the upper layer
individuals in favour of the trees from middle and
lower layer. The diameter distribution on the PRP 2 is
approximating the required model at most. The individuals
from the lower layer (diameter classes 2 and 4
cm) arc moderately deficient. The PRP 3 is covered
with the youngest stand. A part of the plot is in the
growth phase of thicket and small pole-stage stand.
This homogenous part represents an unstable element
which requires a silvicultural measure. The intervention
should reduce the stem number and differentiate
the stand.


According to obtained data the basal area should
range from 35 to 40 m2 ha 1 by the given structure. The
growing stock has quite different values on particular
research plots (from 268.43 m3 ha 1 to 444.22 m ha 1).
Considering the stand structure as well as


the influence of outliers (e.g. one stem with dbh of
88 cm on PRP 1 increases the total growing stock for


more than 80 m3 ha"1) it is possible to propose the optimum
growing stock on the level from 300 to 400 m3
ha"´, according to the production conditions of the site.
The defoliation grade ranges under 10 % on all plots
and on the basis of this characteristic the stands can be
evaluated as vital and healthy.


The analysis of tree frequencies according to the
layers shows that approximately 3/4 of all individuals
are concentrated in the lower layer, whilst other 25 %
are divided by different ratio into middle and upper
layer. After the evaluation of stand structure on particular
plots the most appropriate ratio of the layers seems
to be the model: 75 % lower layer, 15 % middle layer
and 10 % upper layer. The model concerns the ratio according
to the stem number, in the case of area ratio
each layer should take roughly the same area (i.e. 1/3
of the stand area).


Coarse woody debris (CWD) is an important component
of a high-elevation forest especially from the
viewpoint of its regeneration processes. Table 2 sums
up the results of the stand volume divided according to
vital, standing and lying dead trees. The volume of lying
deadwood ranges from 43.56 m3 ha"´ on PRP 2 to


71.67 m3 ha"1 on PRP 1 in absolute values.
Besides the total amount of CWD the ratio between
particular grades of decomposition is important, above
all the volume of CWD in the 2nd and 3rd grade of decomposition,
because only the logs in this state provide
appropriate conditions for the development of natural
regeneration. From this point of view the best conditions
are on the PRP 1 where almost all CWD is at least
partly rotted and creates a convenient seedbed for spruce
seedlings. From given data it is difficult to quantify
the sufficient amount of CWD but it is possible to say
the worse are the conditions for the natural regeneration
in the stand, the higher should be the amount of
CWD. The distribution of the whole biomass on the
observed plots is following: approximately 80-85 %
fall on vital trees, the rest is divided in ratio 1:2 between
standing and lying deadwood. Different situation
is only on PRP 3, where a quite high number of standing
deadwood has been observed.


REGENERATION PROCESSES


Two tree species are present in the natural regeneration
on all research plots - Norway spruce {Picea abies


IL.I Karst.) and rowan (Sorbus aucuparia L.). Although
the tables 3 and 4 show also the total number of
individuals (spruce + rowan), this values as well as the
comparison of the numbers according to the tree species
have only an orientation meaning. The first reason
is, that although rowan is an important and permanent